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Following the appearance of a black cloud resembling a mountain, Godzilla emerges from Mount Fuji and begins a destructive rampage despite having become tolerant of humans within the past years. Godzilla's ally, Anguirus, confronts Godzilla, only to be nearly killed and forced to retreat. Keisuke arrives shortly after to check on Masahiko and Miyajima. Godzilla's rampage continues until another (and the real) Godzilla emerges and battles it. During the fight, the first Godzilla is revealed to be an imposter called Mechagodzilla, a massive robot armed with advanced weaponry made of the same strange metal, later revealed to be space titanium. Godzilla is severely wounded, but inflicts some damage on Mechagodzilla before both monsters retreat. Miyajima hypothesizes, based on Mechagodzilla's advanced technology and composition, that the robot is an alien superweapon.
Masahiko, Miyajima and his daughter Ikuko explore the cave where the space titanium was first found, but are captured by ape-like aliens from the Third Planet of the Black Hole, who plan to use Mechagodzilla to conquer Earth. Their leader, Kuronuma, forces Miyajima to repair the robot. While Saeko checks into a hotel and guards the statue, Keisuke leaves to meet Masahiko at the caves, only to encounter the reporter, who reveals his real name is Nanbara and that he is an Interpol agent who has been tracking the aliens. After Nanbara and Keisuke infiltrate the alien base and free the prisoners, Keisuke and Ikuko leave to pick up Saeko and the statue while Miyajima, Nanbara, and Masahiko stay behind, only to be recaptured by Kuronuma.
The next morning, a lunar eclipse results in a red moon and a mirage of the sun rising in the west. Seeing this, the team realizes they have to awaken King Caesar. They meet with the priestess and her grandfather and place the statue in the temple, revealing King Caesar's resting place. As Kuronuma dispatches Mechagodzilla, the priestess sings to awaken King Caesar and Godzilla appears shortly afterward. The two monsters join forces to fight Mechagodzilla. When the robot tries to escape, Godzilla uses its stored electricity to create an electromagnetic field to attract Mechagodzilla before tearing off its head; causing it to explode. While the mortified aliens are distracted, Nanbara and the others free themselves, kill their captors, and sabotage the base, fleeing as it explodes. With the enemy defeated, Godzilla heads out to sea and King Caesar returns to its resting place while the humans rejoice.
Meanwhile, Bernie Hayes, an employee of the Apex Cybernetics corporation and host of a Titan conspiracy theory podcast known as the Titan Truth Podcast, exfiltrates information suggesting sinister activities at a Pensacola Apex facility, but as he finished downloading the files, Godzilla arrives and unexpectedly attacks. During Godzilla's rampage, Bernie discovers a strange piece of technology resembling a giant robotic eye among the ruins of the facility. In the aftermath of the attack, public opinion has turned against Godzilla, who is no longer viewed as humanity's savior. Walter Simmons, Apex's founder and CEO, promises mankind that he and his company will find a way to defend the world from Godzilla. Meanwhile, Madison Russell, now 17 years old and a fan of Bernie's podcast, sees the news about Godzilla's recent attack at her school and is worried. After school, she tries unsuccessfully to talk sense into her widowed father Mark (who is acting as Monarch's director), regarding a possible lead on the cause of Godzilla's rampage, but he dismisses everything she says as he only replies that creatures, just like people, can change. Later, Madison enlists the help of her friend Josh Valentine so they can investigate the one responsible for causing Godzilla's rampage, have his name cleared, prove the Titan's innocence and figure out the secrets of Apex.
Meanwhile, Madison and Josh find Bernie, who recognizes Madison as Mark's daughter and becomes part of their investigation, believing Apex had something to do with Godzilla's change in behavior. Together, they sneak into the wrecked Apex base, only to find that the "eye" Bernie had described and seen earlier is now gone. As they discover a secret facility deep underground, the group are inadvertently locked and transported in an antigravity transport vehicle carrying Skullcrawler eggs all the way through an underground tunnel to Hong Kong. There, the three disembark and find themselves inside a huge dark chamber, but stumble upon Mechagodzilla, a mechanized counterpart of Godzilla being controlled by the late Dr. Ishiro Serizawa's son and Walter's chief technology officer Ren Serizawa. To test Mechagodzilla's might against the Titans as soon as the mecha is activated, Walter unleashes a large Skullcrawler codenamed "Number 10" into the chamber. The beast targets Madison, Josh and Bernie and is on the verge of attacking them when it is suddenly grabbed and restrained by Mechagodzilla, who cleaves the Skullcrawler in half using a powerful red energy beam, allowing Madison, Josh and Bernie to seek shelter in a small hatch. However, the test is cut short when the mecha powers down immediately afterwards due to limitations of the power supply. It is then revealed that Walter intends to harness the power of the Hollow Earth's energy to overcome Mechagodzilla's limitations. Madison realized that Mechagodzilla is the reason why Godzilla attacked in the first place: Apex activated the mecha which sent out signals that provoked him into becoming hostile so they would manipulate mankind that he has turned against them while also secretly planning to get rid of him and all the Titans so that humans can become the Earth's dominant species. While trying to find an exit, Madison and the others suddenly come upon a room where Ren is sitting inside the skull of Ghidorah's severed head, which has now turned into a biological supercomputer and cockpit for Ren to control Mechagodzilla through radio waves after Apex acquired it from Alan Jonah since Godzilla previously tore it off during his battle against Ghidorah off the coast of Isla de Mara five years ago.
Back in the Apex base, Walter excitedly orders Ren to send out Mechagodzilla to kill Godzilla. Though Ren warns him against using the new Hollow Earth energy source without testing, Walter refuses to listen and coldly orders him to activate the mecha. At the same time, Bernie, Madison, and Josh are caught by security for infiltrating Apex headquarters and taken to Walter. There, Madison accuses Walter of provoking Godzilla into war and for causing all of this in the first place, though he confidently explains that he accepted the responsibility for giving mankind a chance to fight back against Godzilla and the Titans. Unbeknownst to Walter as his back is turned while he monologues about his creation, Mechagodzilla suddenly goes haywire under the influence of both the energy source and Ghidorah's neural networks, resulting in the robot killing Walter with a swipe of its hand and Ren getting electrocuted in the process. Now possessed by Ghidorah's consciousness, Mechagodzilla breaks free from the Apex facility underneath Victoria Peak Mountain and rampages through the city, only to be confronted by Godzilla, who senses the presence of Ghidorah inside the machine. Quickly coming into contact with Godzilla, Mechagodzilla engages its organic counterpart in battle.
While Josh and Bernie unsuccessfully try to deactivate the rampaging robot remotely and Madison tries to call up Mark for help, Godzilla and Mechagodzilla battle, at one point both fire their energy beams at each other which lock together, only for Mechagodzilla to gain the upper hand, knocking Godzilla over and burning him due to the fact that he has already grown weary from his earlier battle against Kong. While Mechagodzilla continues to overpower Godzilla, Nathan, Ilene, and Jia approach an unconscious Kong, where Jia senses that he is dying as his heartbeat is starting to slowly shut down. Ilene explains that they can't produce a charge big enough to restart Kong's heart, but Nathan, having remembered what Maia said earlier about the HEAVs producing a charge powerful enough to "light up Las Vegas for a week", flies the remaining HEAV onto Kong's chest and sets it to self-destruct, detonating the craft and releasing a huge electrical charge which instantly revives Kong. After Kong regains consciousness, he awakens in time to witness Godzilla getting beaten up by Mechagodzilla before reuniting with Jia, who convinces Kong that Godzilla isn't really his enemy. Though Kong doesn't believe her at first as he continues to watch Mechagodzilla overwhelm Godzilla, Jia reveals the truth to him that the robot is the "real" enemy, not Godzilla. After a moment, Kong, realizing that he made a mistake, finally accepts and, after fixing his dislocated arm, heads into battle to help Godzilla, who is now at Mechagodzilla's mercy.
Just when all hope seems lost as Mechagodzilla prepares to fire its red energy beam down Godzilla's throat, Kong leaps onto the robot's back as it was about to deliver the killing blow and redirects the beam into the air, saving Godzilla's life in the process. Surprised that Kong saved his life despite having been beaten to the brink of death earlier and sensing an ally in him upon realizing Kong is not the enemy he thought he was, Godzilla helps his former rival fight against Mechagodzilla. Working together as a team, the two Titans turn the tide, but the robot still proves too strong and is able to overpower them. Seeing his axe among the wreckage after losing it during his earlier battle against Godzilla, Kong reclaims it and lands several blows on Mechagodzilla, but his weapon, having lost its charge, is ineffective against the robot, who attempts to fatally impale Kong with its tail drill. Meanwhile, Bernie, realizing there is nothing he and the others can do to help the two Titans, pulls out a flask of liquor, suggesting that they all have a drink. However, Josh, suddenly having an idea, snatches the flask out of Bernie's hand and short-circuits Mechagodzilla's controls by pouring the contents of the drink all over the control panel, momentarily interrupting the robot in order to give Godzilla and Kong the opportunity to recover. Seeing Kong holding his axe as he struggles to hold his own against Mechagodzilla, Godzilla supercharges the weapon by blasting it with his atomic breath, giving Kong a second wind and allowing him to dismember Mechagodzilla limb from limb before tearing its head off. With the threat eliminated, Kong holds up his trophy with a mighty bellow of triumph before sitting down to rest. 2b1af7f3a8
